Cats may have a reputation for independence, but emerging research suggests we share a unique connection with them – fuelled by brain chemistry.

The main chemical involved is oxytocin, often called the love hormone. It’s the same neurochemical that surges when a mother cradles her baby or when friends hug, fostering trust and affection. And now studies are showing oxytocin is important for cat-human bonding too.

Oxytocin plays a central role in social bonding, trust and stress regulation in many animals, including humans. One 2005 experiment showed that oxytocin made human volunteers significantly more willing to trust others in financial games.

Oxytocin also has calming effects in humans and animals, as it suppresses the stress hormone cortisol and activates the parasympathetic nervous system (the rest and digest system) to help the body relax.

Scientists have long known that friendly interactions trigger oxytocin release in both dogs and their owners, creating a mutual feedback loop of bonding. Until recently, though, not much was known about its effect in cats.

Cats are more subtle in showing affection. Yet their owners often report the same warm feelings of companionship and stress relief that dog owners do – and studies are increasingly backing these reports up. Researchers in Japan, for example, reported in 2021 that brief petting sessions with their cats boosted oxytocin levels in many owners.

In that study, women interacted with their cats for a few minutes while scientists measured the owners’ hormone levels. The results suggested that friendly contact (stroking the cat, talking in a gentle tone) was linked to elevated oxytocin in the humans’ saliva, compared with a quiet resting period without their cat.

Many people find petting a purring cat is soothing, and research indicates it’s not just because of the soft fur. The act of petting and even the sound of purring can trigger oxytocin release in our brains. One 2002 study found this oxytocin rush from gentle cat contact helps lower cortisol (our stress hormone), which in turn can reduce blood pressure and even pain.

When is oxytocin released between cats and humans?

Research is pinpointing specific moments that cause the release of this hormone in our cross-species friendship. Gentle physical contact seems to be a prime trigger for cats.

A February 2025 study found that when owners engaged in relaxed petting, cuddling or cradling of their cats, the owners’ oxytocin tended to rise, and so did the cats’ – if the interaction was not forced on the animal.

The researchers monitored oxytocin in cats during 15 minutes of play and cuddling at home with their owner. Securely attached cats who initiated contact such as lap-sitting or nudging showed an oxytocin surge. The more time they spent close to their humans, the greater the boost.

What about less-cuddly felines? The same study noted different patterns in cats with more anxious or aloof attachment styles. Avoidant cats (those who kept their distance) showed no significant oxytocin change, while cats who were anxious (constantly seeking their owner but easily overwhelmed by handling) had high oxytocin to begin with.

Oxytocin of avoidant and anxious cats was found to drop after a forced cuddle. When interactions respect the cat’s comfort, the oxytocin flows – but when a cat feels cornered, the bonding hormone is elusive.

Maybe humans could learn something from their feline friends on managing attachment styles. The key to bonding with a cat is understanding how they communicate.

Unlike dogs, cats don’t rely on prolonged eye contact to bond. Instead, they use more understated signals. The most well known is the slow blink. It’s a feline smile, signalling safety and trust.

Purring also plays a role in bonding with people. The low-frequency rumble of a cat’s purr has been linked not only to healing in cats themselves, but also to calming effects in humans. Listening to purring can lower heart rate and blood pressure; oxytocin mediates these benefits.

The companionship of a cat, reinforced by all those little oxytocin boosts from daily interactions, can serve as a buffer against anxiety and depression – in some cases providing comfort on par with human social support.

Are cats just less loving than dogs?

It’s true that studies generally find stronger oxytocin responses in dog–human interactions. In one widely discussed 2016 experiment, scientists measured oxytocin in pets and owners before and after ten minutes of play. Dogs showed an average 57% spike in oxytocin levels after playtime, whereas cats showed about a 12% increase.

In humans, oxytocin levels rise during meaningful social interactions. Studies show that contact with a loved one produces stronger oxytocin responses than contact with strangers. So, a happy dog greeting is akin to that rush of seeing your child or partner.

Dogs, being pack animals domesticated for constant human companionship, are almost hard-wired to seek eye contact, petting and approval from us – behaviour that stimulates oxytocin release in both parties. Cats, however, evolved from more solitary hunters which didn’t need overt social gestures to survive. So, they may not display oxytocin-fueled behaviour as readily or consistently. Instead, cats may reserve their oxytocin-releasing behaviour for when they truly feel safe.

A cat’s trust isn’t automatic; it must be earned. But once given, it is reinforced by the same chemical that bonds human parents, partners and friends.

So, next time your cat blinks slowly from across the sofa or climbs on to your lap for a purr-filled cuddle, know that something invisible is happening too: oxytocin is rising in both your brains, deepening the trust and soothing the stress of daily life. Cats, in their own way, have tapped into the ancient biology of love.

Laura Elin Pigott, Senior Lecturer in Neurosciences and Neurorehabilitation, Course Leader in the College of Health and Life Sciences, London South Bank University

This article is republished from The Conversation under a Creative Commons license. Read the original article.

While often miscast as mysterious or hard to understand, cats are actually excellent communicators. In fact, in free-ranging cat colonies, physical fights are kept to a minimum through clever use of body posturing, scent exchange and vocalisations.

Cats have also adapted their communication for humans. For instance, adult cats don’t usually meow at each other. But when around people, cats meow a lot, suggesting they have adapted this vocalisation for communicating with humans.

And it’s not just the meow. Cats have a wide vocal repertoire for conveying different meanings, even for specific people. Bonded cats and humans often develop their own communication repertoires, similar to having a unique dialect.

Cats can understand human communication too. Studies show cats know their own names and the names of their companions, and can recognise human emotions, even changing their own behaviour in response.

Despite all this, humans still routinely misunderstand cats. Our new study, published in Frontiers in Ethology, shows just how little people understand the cues cats give. Try the quiz below to see how well you speak cat.

What we did

We asked 368 Australian participants to watch videos of human–cat play interactions. But not all the videos were “play” for the cat. Only half of the cats were playing, while the other half were actually showing signs they didn’t want to play, or were feeling stressed by the interaction.

After each video, participants were asked if they thought the interaction was overall positive or negative for the cat, based only on the cat’s behaviour. They were then asked how they would interact with the cat in the video they had just seen.

How well do you speak cat?

Watch the short videos below and decide: is the cat feeling positive or negative about the interaction? Remember to base your answers only on the cat’s behaviour.

What did our study find?

Results showed that participants struggled to recognise negative cues indicating discomfort or stress in cats.

For videos of cats who weren’t playing and were showing subtle negative cues (such as sudden tension in the body or avoiding touch), participants only recognised the negative cues about as well as chance (48.7%).

Even when participants watched videos of cats showing overt negative cues like hissing, biting or trying to escape, they still incorrectly categorised these as positive 25% of the time.

Recognising when a cat is stressed is only the first step. We also need to know how to respond to these cues.

Even when participants did successfully recognise negative cues, they often chose to engage with the cat in ways that would cause more stress and increase the risk of human injury, such as stroking, belly rubbing and playing with hands.

Stress is unhealthy

Stress can have serious consequences. Cats who experience regular or prolonged stress (including from unwanted interactions like those in the negative videos) are at higher risk of heath issues such as bladder inflamation.

They’re also more likely to develop behaviours people find problematic, such as increased aggression or urinating outside of the litter tray. In turn, these behaviours increase risk of the cat being euthanised or rehomed.

Cat stress is bad for humans, too. If a person doesn’t heed early warning signs, the cat may bite or scratch, depositing bacteriaand microorganisms deep into the skin. Rapid infection follows 30%–50% of cat bites. If not treated promptly, it can lead to serious complications including sepsis, chronic health issues and even death. Cat bites and scratches can also transmit zoonotic diseases such as cat scratch disease.

How to play safely with cats

Watch for early warning signs a cat isn’t enjoying themselves and stop if you notice any. By the time cues are obvious, cats are already experiencing distress.

Early warnings include turning away, dodging or blocking attempts to touch, flinching, body tension, ears back or to the side, lip/nose licking and tail thrashing, slapping or tucking.

Touch

Avoid sensitive areas such as the belly, paws or the base of the tail. Cats prefer to be touched on the head and neck.

Avoid using hands to play. It teaches cats that hands are toys, and increases the risk of accidental injury. Instead, use toys that keep your face and hands away, such as a wand toy with a long handle.

Tail

Tail movements aren’t always a negative sign – they just mean the cat is emotionally stimulated and that could be from stress or excitement. Cats also use their tails for balance. So it’s best to consider the tail in combination with the whole body and the context.

Changes in tail movements can also give important clues to the cat’s mood. Generally, the bigger the movement, the more intense the feeling. So, if the movements start to get bigger or faster during play, or if a tail goes from relaxed to swishing when you touch, that might be a sign to back off.

Ears

Cats’ ears are like antennas that swivel and adjust to pinpoint sound, but they can also give us a clue to how they are feeling. If the ears move for a moment and then return to a relaxed position, that usually means they’re listening to the world around them. If the ears remain flattened and back, that’s a sign of distress.

Vocalisations

Trilling and chirruping both suggest a playful cat, while hissing, growling and yowling all indicate stress. Purring might seem positive but may indicate a cat is stressed and trying to self soothe.

Let them be

When early warnings don’t work, cats may show overt signs such as hissing, growling, trembling, hiding and, ultimately, biting or scratching.

If you notice warning signs, give the cat plenty of space. When stressed, cats don’t like being touched or having people too close. If the cat comes back and re-initiates contact, that’s a good sign they’re comfortable, but keep watch for warning signs returning.

If you pay attention to your cat’s behaviour and give them space when they need it, with a bit of practice you might just become fluent in cat.

Julia Henning, PhD Candidate in Feline Behaviour, School of Animal and Veterinary Science, University of Adelaide

This article is republished from The Conversation under a Creative Commons license. Read the original article.

Twitter’s been on fire with people amazed by cats that seem compelled to park themselves in squares of tape marked out on the floor. These felines appear powerless to resist the call of the #CatSquare.

This social media fascination is a variation on a question I heard over and over as a panelist on Animal Planet’s “America’s Cutest Pets” series. I was asked to watch video after video of cats climbing into cardboard boxes, suitcases, sinks, plastic storage bins, cupboards and even wide-necked flower vases.

“That’s so cute … but why do you think she does that?” was always the question. It was as if each climbing or squeezing incident had a completely different explanation.

It did not. It’s just a fact of life that cats like to squeeze into small spaces where they feel much safer and more secure. Instead of being exposed to the clamor and possible danger of wide open spaces, cats prefer to huddle in smaller, more clearly delineated areas.

When young, they used to snuggle with their mom and litter mates, feeling the warmth and soothing contact. Think of it as a kind of swaddling behavior. The close contact with the box’s interior, we believe, releases endorphins – nature’s own morphine-like substances – causing pleasure and reducing stress.

Along with Temple Grandin, I researched the comforting effect of “lateral side pressure.” We found that the drug naltrexone, which counteracts endorphins, reversed the soporific effect of gentle squeezing of pigs. Hugs, anyone?

Also remember that cats make nests – small, discrete areas where mother cats give birth and provide sanctuary for their kittens. Note that no behavior is entirely unique to any one particular sex, be they neutered or not. Small spaces are in cats’ behavioral repertoire and are generally good (except for the cat carrier, of course, which has negative connotations – like car rides or a visit to the vet).

One variation on this theme occurs when the box is so shallow that it does not provide all the creature comforts it might.

Or then again, the box may have no walls at all but simply be a representation of a box – say a taped-in square on the ground. This virtual box is not as good as the real thing but is at least a representation of what might be – if only there was a real square box to nestle in.

This virtual box may provide some misplaced sense of security and psychosomatic comfort.

The cats-in-boxes issue was put to the test by Dutch researchers who gave shelter cats boxes as retreats. According to the study, cats with boxes adapted to their new environment more quickly compared to a control group without boxes: The conclusion was that the cats with boxes were less stressed because they had a cardboard hidey-hole to hunker down in.

Let this be a lesson to all cat people – cats need boxes or other vessels for environmental enrichment purposes. Hidey-holes in elevated locations are even better: Being high up provides security and a birds’s-eye view of the world, so to speak.

Without a real box, a square on the ground may be the next best thing for a cat, though it’s a poor substitute for the real thing. Whether a shoe box, shopping bag or a square on the ground, it probably gives a cat a sense of security that open space just can’t provide.

Nicholas Dodman, Professor Emeritus of Behavioral Pharmacology and Animal Behavior, Cummings School of Veterinary Medicine, Tufts University

This article is republished from The Conversation under a Creative Commons license. Read the original article.

For the sharpest minds in show business, there’s always another hustle.

Take Joe Exotic, whose 2020 conviction for a murder-for-hire plot and violations of the Endangered Species Act hasn’t kept the eccentric tiger trainer out of the headlines.

Since beginning his 21-year sentence, the “Tiger King” star has started a cannabis brand, hawked digital art and begun work on an album tentatively titled “Jungle Rhapsody: A Tiger King Experience.” His most recent gambit involves selling personal phone calls from his cell – “What better Valentines gift could you ever get your loved one,” he posted on Instagram in January 2026.

But before Joe Exotic, there was Mabel Stark. Often described as America’s most famous female tiger trainer, the Tiger Queen was renowned for her pluck and charisma.

While researching Caxton Printers, the publisher of Stark’s autobiography, I came across unpublished archival material about Stark’s long career training animals. Like Joe Exotic, Stark had a knack for the spotlight. But even more impressively, she did it under the scrutiny of being a woman in a male-dominated world, while caring for her animals with love rather than fear.

From nurse to tiger trainer

Born Mary Ann Haynie in 1888 or 1889 – the exact year has always been a mystery – Stark grew up in Princeton, Kentucky. When she was 8 years old, she attended her first circus, where she was awed by the performances of trained animals.

Two decades would pass before she got a chance to try her hand at animal training.

Taking a vacation in California from her job as a nurse, Stark met Al Sands, manager of the Al G. Barnes Circus. After learning of her interest in training animals, he hired her on the spot.

Stark started by riding horses and training goats. It would take several years before she started working with tigers. But once she did, her career took off.

Crowds gathered to watch the “Tiger Girl” wrestle with big cats and wow audiences by commanding a dozen tigers at a time to follow her lead. Her wrestling act with her favorite tiger, Rajah – in which the duo would roll three or four times on the ground – became one of the best-known cat acts in the U.S.

She leveraged that success to join the Ringling Circus – the largest circus in the U.S. – for twice the pay.

As her popularity grew, Stark collaborated with screenwriter Gertrude Orr to write her life’s story.

“Hold That Tiger” hit bookstores in 1938. Caxton Printers, a small publishing company in rural Idaho, issued the book and marketed it primarily to young readers. It proved popular, selling well enough to warrant multiple reprintings.

Known for giving a voice to first-time writers and authors from underrepresented groups, Caxton Printers found a niche market for circus-related titles. It also published books about Stark’s first employer, Al Barnes, as well as the Ringling Brothers and renowned lion trainer Louis Roth, who also happened to be one of Stark’s ex-husbands.

Female power in the ring

Stark was acutely aware of the path she was paving.

“I deliberately chose a field in which no other woman had specialized,” she wrote in her autobiography.

The conventional wisdom at the time, she added, was that “tigers were considered too dangerous for a woman to handle.”

Stark’s willingness to defy convention mattered. As circus historian Janet M. Davis noted, “circus women’s performances celebrated female power” and represented “a startling alternative to contemporary social norms.”

In early-20th-century American life, women might not have been able to vote or to serve on juries in most states, but in the ring, they commanded the audience’s attention riding bareback on horses, displaying strength and stamina, and performing gravity-defying acrobatic feats.

Stark’s schedule was relentless. She performed almost daily with traveling circuses, and she continually refined her act. In 1938, she worked with both tigers and lions at the same time, a first for a female trainer. She made history again working with 12 tigers in one cage.

Whether it was due to the demands of her schedule or her preference for her cats, Stark’s relationships with men rarely worked out.

Over the course of her life, Stark married four times, three of which ended in divorce.

“I love these big cats as a mother loves her children,” she admitted to a friend. But “with husbands I was never happy.”

‘An animal trainer can’t have nerves’

Stark, aware of other trainers’ abusive behavior toward their tigers, took a different route.

“Kindness and patience are the biggest factors in training. … Trainers who try to beat animals into submission always get into trouble,” she said.

Yet her trade was not without danger.

“An animal trainer can’t have nerves. I haven’t had any since I gave up nursing,” she said in a 1922 New York Times interview. “They may be planting violets on me tomorrow, but while I have my health and strength, I’d rather take care of 10 tigers than a sick person.”

Stark had several serious accidents. Perhaps the worst was in 1928: After a circus train arrived late, Stark started her act without realizing her tigers hadn’t been fed for 24 hours. Two famished tigers attacked Stark after she fell in mud.

“As I lay there, helpless,” she wrote, “I wondered into how many pieces I would be torn, and how long it would take for the other tigers, growling and snarling restlessly on their seats, to finish me.” She suffered multiple broken bones, nearly lost her leg and required 300 stitches.

Another incident took place in 1950, when a tiger mauled her as she reached for its cub. Doctors initially thought they would have to amputate her arm but managed to save it.

Despite these close calls with her tigers, Stark maintained that “I am not afraid. I like the challenge of their roaring defiance.”

The stark reality

Stark toured with circuses until the late 1940s, when she was hired by Jungleland, a zoo located outside of Los Angeles.

Save for the three-and-a-half years she lived in Japan touring with her wild cat act, she spent the last 20 years of her career at Jungleland.

Stark never stopped drawing crowds to her show, nor did she shy away from the spotlight. She even appeared on the game show “What’s My Line?” in 1961 as a contestant whose profession the panel had to guess.

“Each year has left scars on my body, but it has also brought a full measure of happiness,” she recalled. Stark worked at Jungleland until she was fired in 1967 after the park’s insurance company stopped covering her. Being away from her tigers devastated her, and she died by suicide just months later on April 20, 1968, at her home in Thousand Oaks.

The concluding paragraph of Stark’s autobiography anticipates the end of her life:

“The chute door opens as I crack my whip and shout, ‘Let them come!’ Out slink the striped cats, snarling and roaring, leaping at each other or at me. It’s a matchless thrill, and life without it is not worth while to me.”

Alessandro Meregaglia, Associate Professor and Archivist, Boise State University

This article is republished from The Conversation under a Creative Commons license. Read the original article.

Larry, the Downing Street cat, has had to adapt to seven different prime ministers moving into No 10 over the past decade. Yet while the nation’s attention is fixed on the departing and incoming political leaders, who is thinking about the effect these upheavals might have on one of the property’s most enduring residents?

Cats thrive on predictability and the consistency of their environment is important for their welfare. On the whole cats prefer an unchanging environment. Some cats cope with change better than others. Many cats are distressed by disturbances such as moving house, changes in familiar routines, people moving into or leaving the household, or even changes to the scent of their home environment.

Unlike dogs, cats aren’t obligate social species, therefore may not have such a strong proclivity to form social attachments. However, in certain situations domestic cats can be found living in social groups, particularly in environments with a plentiful food supply.

Some cats enjoy the company of people, while others prefer a more solitary life. This is determined partly by their early experiences. A young kitten that learns in the early weeks of their life that contact with people is associated with nice experiences is more likely to seek social interactions with people as an adult. Kittens raised by mothers who show friendly behaviour towards people can learn to value human company through the process of social learning.

Genetics also influence a cats friendliness towards people. Researchers in a 1995 study found that the kittens of friendlier fathers were more likely to show friendly behaviour towards unfamiliar people even if they weren’t socialised as a new kitten.

Some cats appear to suffer when separated from their caregivers, although some researchers question whether this can be described as a secure attachment (one that is necessary to maintain feelings of safety and security in threatening situations).

Researchers have identified five different forms of cat-owner relationship, based on the level of emotional investment of the owner, the way the cat relates to others, their aloofness and their need to be in close proximity to their owner. These types range from open relationships (characterised by a lightly emotionally invested owner and an avoidant cat) to casual relationships, co-dependence and friendship.

Although the Starmer family has moved out and the Burnhams have taken up residence, Larry’s main caretakers are civil servants, so his day-to-day care won’t change. Larry’s behaviour in recent days (strutting along Downing Street, nonplussed by the comings and goings and embarking on a successful hunting expedition) suggests that he takes such changes in cohabitants in his stride.

Other animals

The new prime minister has a dog called Axel, and although cats and dogs can live together harmoniously and even form close bonds, integrating Axel into Larry’s home might cause tensions if not handled well. However, not even the prime minster would dare to disturb Larry’s position as chief mouser. Andy Burnham has told the BBC he’s not bringing Axel to London with him because he doesn’t think the two animals will get on.

If he changes his mind there are a few things he could do to help Axel and Larry get along. Introducing a dog into a household with a cat should be done gradually, with the aim to prevent any chasing or aggression between the pets.

Before the dog arrives, settle the cat into a safe space, which the dog won’t have access to. This might be a bedroom, bathroom or any part of the house that the cat likes to relax in, ensuring the cat has access to everything they need such as places to hide, litter trays, food and water.

Scent is incredibly important to both cats and dogs, so getting them accustomed to each other’s aroma in the home helps to smooth the introduction. This can be done by introducing bedding or toys used by the other pet, or gently wiping a damp cloth across the body of the pet to pick up their scent.

It’s vital that the first meeting goes well and there are a few things that will help to ensure they get off on the right foot. Make sure the dog isn’t full of energy by exercising them before the meeting. Start with a barrier such as a baby gate between them.

When they first meet give both pets plenty of their favourite food treats, to reward calm behaviour and so that they associate the presence of other with something nice happening.

Signs of stress

Cats can be difficult to read but there are known warning signs of stress in cats. More obvious stress behaviour includes hissing, growling, swiping or puffed up fur. But indicators also include the cat running away or hiding and refusing treats they normally avoid. Flattened ears, dilated pupils, tense body posture and a flicking tail are all signals a cat is feeling tense.

Larry seems like quite an independent confident cat if reports “he has the run of the house” are anything to go by, so it’s likely that he will soon teach the new inhabitants of No 10 how he likes things done.

Emily Blackwell, Senior Lecturer in Animal Behaviour and Welfare, University of Bristol

This article is republished from The Conversation under a Creative Commons license. Read the original article.

It’s pretty commonly known, and not very startling, that free-ranging cats eat birds and small rodents. But the degree to which they eat insects might surprise you.

Wearebiologists who for many years have been trying to figure out what feral or outdoor-roaming pet cats eat outside.

When domesticated cats – Felis catus – live freely in the wild or are allowed to hunt outside the homes where their owners live, they are an invasive species, which live in every ecosystem of the world except the continent of Antarctica. We wanted to know all of the species they eat – and to what degree free-ranging cats are eating endangered or threatened species.

Examining reams of research

Over the past two decades, we have evaluated hundreds of scientific findings, including searching through Google Scholar and Web of Science using the keywords “cat predation,” “feral cat,” “cat diet” and “Felis catus.” For each item we found, whether peer-reviewed or not, we evaluated whether it contained conclusive evidence of cat diet or predation. We also reviewed each one’s reference section for additional unique articles or databases pertaining to cat diet and predation, and included those in our search.

Overall, we identified 533 unique publications – books, journal articles, theses and agency reports – that reported specific animal species consumed by cats. Cats’ plant-eating habits are occasionally, but haphazardly, noted in studies, so we did not include them in our analysis.

Our initial work focused on an overall assessment of what free-ranging cats eat around the world. Published in 2023, this paper analyzed the 533 studies on cat diet or predation events published over more than a century and found that cats ate nearly 2,100 different species of animals, including invertebrates.

Of those 2,100, the International Union for Conservation of Nature’s Red List of Threatened Species listed 347 as “near threatened,” “vulnerable,” “endangered,” “critically endangered” or “extinct” in 2023. Some of the species went extinct during the many decades covered by the data.

Insects and the like

Most of the species cats ate were vertebrates – mostly birds, followed by mammals and reptiles. But the data also indicated that at least 7% of the species cats eat are insects and other invertebrates, particularly beetles, and less frequently crustaceans, arachnids, centipedes, snails and slugs, and millipedes.

Many of the cat studies we reviewed did not report on how many individuals of a given species cats ate, so it was unclear what the total amount of insects was or how many calories cats are deriving from insects.

Invertebrates make up more than 70% of all terrestrial animal species and are important pollinators, predators and herbivores in virtually every nonmarine ecosystem. Many invertebrates are in decline globally due to urbanization, habitat destruction, increases in both light and pesticide pollution, and climate change. So we dug deeper into the data to understand what invertebrates cats are eating.

While a little more than one-third of all the studies we analyzed included invertebrates as part of cats’ diet, most of those failed to identify specific species of invertebrates. But we were able to find identifications of 148 invertebrate species.

Of those, two are considered endangered by the International Union for Conservation of Nature: the Aldabran grasshopper (Pternoscirtus aldabrae) in Seychelles and the Tasmanian giant freshwater crayfish (Astacopsis gouldi), which can grow up to 13 pounds (6 kilograms). Two others are considered vulnerable: wētāpunga (Deinacrida heteracantha), an insect native to New Zealand that can be about the size of a mouse, and the common yabby (Cherax destructor), a freshwater crayfish native to southeastern Australia. One other, the Canary Islands horned beetle (Arhopalus pinetorum), is listed as “near threatened.”

Effects on populations

We have not found formal research evaluating how cats’ eating habits affect invertebrate populations. And for many species, they are likely not as significant a factor as wide-scale pesticide use.

But it’s possible that cats could be significant contributors to the deaths of rare species or in specific locations.

Cats require a large amount of protein, as much as one-third of their daily diet, and invertebrates are good sources of protein.

In many places, invertebrates provide an easy source of food. Whether in an urban backyard or on a remote island, cats are unlikely to turn a blind eye to available prey. And some cats may find it entertaining to chase, catch and eat insects even if they don’t need their nutrition.

A challenge of researching this question is that many invertebrates are relatively small, which makes direct observation in the field harder and can require more analytical approaches in the lab. And they have soft bodies, without distinguishing characteristics that could be easily recognized in scat or stomach contents.

However, molecular technologies can identify species using trace amounts of DNA left in the environment by animals. Promising new studies are beginning to identify what cats eat by analyzing the DNA found in their stomachs and scat. That research may help explore in even more detail what cats are eating in the outdoors, and how it’s affecting various species and the environment as a whole.

Christopher A. Lepczyk, Professor of Wildlife Biology and Conservation, Auburn University; Daniel Rubinoff, Professor of Entomology, University of Hawaii, and Jean E. Fantle-Lepczyk, Research Fellow, College of Forestry, Wildlife and Environment, Auburn University

This article is republished from The Conversation under a Creative Commons license. Read the original article.

Diane Keaton loved her dog, Reggie.

The award-winning actor, director and real estate entrepreneur frequently posted photos and video clips of the golden retriever on her social media accounts. After she died on Oct. 11, 2025, at 79, some news outlets reported that she left US$5 million of her estimated $100 million estate to her dog.

I’m a law professor who teaches about wills, trusts and other forms of inheritance law. Every semester, I teach my students how they can help clients provide for their pets after death. Because they, like many Americans, love their pets and want to know how to take care of them, this topic always piques their interest.

Writing pets into a will

An estimated 66% of all U.S. households include at least one pet. Many Americans consider their cats, dogs, tortoises or other animals to be part of their family, and their spending on those nonhuman relatives is immense. In 2024, they paid a total of about $152 billion for goods and services to feed and otherwise support their pets.

Taking good care of your pets can go beyond buying them treats and sweaters. It can include leaving clear directions to ensure their needs are met once you’re gone. There are several ways that you can do this.

The first is through your will. You can’t give your pet money directly in your will, because the law says that pets are property, like your books or your dishes.

You can, however, leave a bequest, the technical term for a gift to a person or a cause listed in a will, to someone who will be the animal’s caretaker. That bequest can include directions that the money be spent meeting the pet’s needs.

It’s worth it to also name an alternate or contingent caretaker in case the first person you name does not want to or cannot take on that responsibility, or they die before you or the animals you’ve provided for in the will.

Choupette’s life of luxury

German fashion designer, photographer and creative director Karl Lagerfeld, who died in 2019 at 85, was someone who made the mistake of leaving money directly to his fluffy Birman cat, Choupette. It worked out for Choupette, though.

The cat was, according to several reports, still alive in 2025 and eating meals out of the porcelain bowls that Lagerfeld bought for her. Choupette is cared for at great expense and in the utmost luxury by Françoise Caçote, the designer’s former housekeeper. The cat even had a 13th birthday party at Versailles.

Another pet owner who did right by her pet was the comedian, producer and red carpet interviewer Joan Rivers.

Rivers had two rescue dogs in Manhattan and two more dogs in California when she died in 2014 at age 81. Rivers had made provisions for their care in her will.

Creating pet trusts

If you’d like an arrangement that’s more secure than a will, then you might want to opt for a pet trust, another celebrity favorite. These kinds of trusts were not possible until the 1990s, because pets were not considered true beneficiaries – meaning they couldn’t sue the trustee.

But in the 1990s, states began to change their rules to allow for pet trusts. Today, pet trusts are valid in the whole country, although the rules vary slightly from state to state.

To establish a pet trust, you or a lawyer must draw up a trust document that names two important people: a trustee and a caretaker. The trustee is the person who will manage the money you leave in trust. They will make distributions to the caretaker that you select.

You must also specify how the money is to be spent meeting the animal’s needs and who would get any money that could be left in the trust when the pet dies. Typically, these trusts take effect at the owner’s death, just like other provisions in a will.

Drafting a pet trust can be free, if you use an online template and get no legal guidance. The same thing might cost around $100 if you use an online service such as Legal Zoom that provides directions. More commonly, however, pet trusts are part of a broader estate plan, and costs range depending on how complicated your estate is.

When the rich go overboard

One of the most over-the-top pet trusts came from Leona Helmsley, the New York hotel and real estate mogul known widely as the “Queen of Mean.” She was famous for her pettiness and tough management style and for landing in prison for tax evasion.

When Helmsley died in 2007, she left her dog, a Maltese named Trouble who had reportedly bitten members of her staff, a $12 million trust fund. Most of Helmsley’s estate went to the Helmsley Charitable Trust, but she made individual gifts to several relatives, and the gift to Trouble was larger than any of those.

The grandchildren, upset that Trouble got more money than they did, took the case to court, where the probate judge was less than impressed by Trouble’s luxury lifestyle and knocked down the amount in trust to $2 million. The other $10 million flowed back to her family’s foundation, where the bulk of the estate went in the first place.

Lesson learned: Your dog can have a trust fund, but don’t go overboard.

Bequests for pets can be challenged – in which case it’s up to courts to determines how much they think is reasonable for the pet’s need. In Helmsley’s case, $12 million was found to be excessive. And maybe with good reason. Trouble still had a nice life with fewer millions. The dog died in December 2010 after several years in Sarasota, Florida, at a Helmsley-owned hotel.

Other pet owners who aren’t celebrities have used pet trusts as well, such as Bill Dorris, a Nashville businessman without any human heirs. He left his dog, Lulu, $5 million.

Pet-loving celebrities who loved all the pets

Finally, there’s a lesson to be learned from British fashion designer and icon Alexander McQueen, who was worth £16 million ($21 million) when he died in 2010 at the age of 40. McQueen left £50,000 ($66,000) in a trust for his two bull terriers so that they would be well cared for during the remainder of their lives.

McQueen also included a bequest of £100,000 ($132,000) to the Battersea Dogs and Cats Home in his will to help fund the care of some of the millions of other animals out there that need the basics of food and shelter.

Animal shelters, in the U.K., the United States and other countries, help rescue and protect animals, and these animals need more help than the Choupettes and Troubles of the world.

So, my advice is that you go ahead and create a pet trust for your cat. But don’t forget to give some money in your will – and ideally while you’re alive – to help the vast majority of the millions of companion animals who need new homes every year. None of them have trust funds.

What becomes of Reggie, Keaton’s golden retriever, and her estate remains to be seen. Keaton, who starred in hit movies such as “Annie Hall,” “Reds” and “The First Wives Club,” isn’t the first celebrity to leave millions of dollars to a pet. And it’s unlikely that she will be the last.

Allison Anna Tait, Professor of Law, University of Richmond

This article is republished from The Conversation under a Creative Commons license. Read the original article.